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The 512 file limitation is on the root, not on any folders off the root. An "Audio" folder is technically a subfolder of the root, and not subject to the 512 file/folder limitation.
You can reduce the 512 limit with long filenames, but they would have to be really long, and they would all have to be in the root. Putting files in the root of an SD card makes no sense. Even in a regular drive there are only 2-3 files that need to be in the root. Everything else can/should be in folders. If your needs exceed these limitations, then a Treo with a FAT file system is not what you need; that is what Windows XP with NTFS or UNIX are for. If you are going to have hundreds of MP3 files, put them in a folder called AUDIO and don't worry about it - it should work fine.
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